As discussed recently on hotspot-compiler-dev[1], having a private class with no default constructor can lead to C2 failing to inline, due to the synthetic bridge constructor using a dummy argument of an uninitialized class. This is really a problem in C2, as well as something which could ultimately be resolved by nestmates...
However, there is an easy workaround in adding an empty package-private constructor. In the most recently found case - a microbenchmark stressing MethodHandles.iteratedLoop - adding this to ArrayList$Itr lead to a 2.5-3x speedup. It thus makes sense to do this simple change in this particular case as a quick fix and solve the larger C2 issue separately.
[1] http://mail.openjdk.java.net/pipermail/hotspot-compiler-dev/2016-September/024407.html